Defensive QB's

For the Cowboys it would be Ken Hamlin and if he is not on the field then ma boy  D-Ware

"Sorry seatown, but the MLB is not the person who lines up the front 7 most of the time. "

Wrong, wrong, wrong, wrong.  Who slaps the lineman on the butt to get him to adjust?  The LB's who is talking to the other LB's to get them into their correct position?  The LBs.  Safeties are 15-25 ft behind the line of scrimmage, so if you think a safety can tell the lineman where to line up you are a moron.  LB's are the Shortstops where as the safety is like the centerfielder.  They both are the play callers for there levels.  Safeties have the back four and LB"s have the front seven.  It's a given, you don't even have to know football to see who is adjusting the lineman.

"For the Cowboys it would be Ken Hamlin and if he is not on the field then ma boy  D-Ware"

Ouch, if Ken Hamlin calls your plays your defense is screwed.  That man gets beat deep all the time.  The man was hit in the head with a street sign, you know that had to create some brain damage.

For my Eagles, I'll go with Brian Dawkins. I would say the MLB but like some posters said, the potential for a loss of communication exists with the numerous collisions within the front seven.

I love it a Packer throw away is the captain and signal caller for the Lions